diff --git a/.changeset/fair-radios-knock.md b/.changeset/fair-radios-knock.md
new file mode 100644
index 00000000000..b2c672a98bc
--- /dev/null
+++ b/.changeset/fair-radios-knock.md
@@ -0,0 +1,5 @@
+---
+'@primer/react': patch
+---
+
+Restore Link underline preference to original behavior
diff --git a/.playwright/snapshots/components/BranchName.test.ts-snapshots/BranchName-Default-light-forcedUnderlines-linux.png b/.playwright/snapshots/components/BranchName.test.ts-snapshots/BranchName-Default-light-forcedUnderlines-linux.png
deleted file mode 100644
index 161e327bb67..00000000000
Binary files a/.playwright/snapshots/components/BranchName.test.ts-snapshots/BranchName-Default-light-forcedUnderlines-linux.png and /dev/null differ
diff --git a/.playwright/snapshots/components/Breadcrumbs.test.ts-snapshots/Breadcrumbs-Default-light-forcedUnderlines-linux.png b/.playwright/snapshots/components/Breadcrumbs.test.ts-snapshots/Breadcrumbs-Default-light-forcedUnderlines-linux.png
deleted file mode 100644
index 10b79842e9c..00000000000
Binary files a/.playwright/snapshots/components/Breadcrumbs.test.ts-snapshots/Breadcrumbs-Default-light-forcedUnderlines-linux.png and /dev/null differ
diff --git a/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Default-light-forcedUnderlines-linux.png b/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Default-light-forcedUnderlines-linux.png
deleted file mode 100644
index 0ae25820262..00000000000
Binary files a/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Default-light-forcedUnderlines-linux.png and /dev/null differ
diff --git a/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Muted-light-forcedUnderlines-linux.png b/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Muted-light-forcedUnderlines-linux.png
deleted file mode 100644
index ea2038fd8fb..00000000000
Binary files a/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Muted-light-forcedUnderlines-linux.png and /dev/null differ
diff --git a/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Underline-light-forcedUnderlines-linux.png b/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Underline-light-forcedUnderlines-linux.png
deleted file mode 100644
index 0ae25820262..00000000000
Binary files a/.playwright/snapshots/components/Link.test.ts-snapshots/Link-Underline-light-forcedUnderlines-linux.png and /dev/null differ
diff --git a/.playwright/snapshots/components/LinkButton.test.ts-snapshots/LinkButton-Invisible-light-forcedUnderlines-linux.png b/.playwright/snapshots/components/LinkButton.test.ts-snapshots/LinkButton-Invisible-light-forcedUnderlines-linux.png
deleted file mode 100644
index 9984199a841..00000000000
Binary files a/.playwright/snapshots/components/LinkButton.test.ts-snapshots/LinkButton-Invisible-light-forcedUnderlines-linux.png and /dev/null differ
diff --git a/e2e/components/BranchName.test.ts b/e2e/components/BranchName.test.ts
index 0da26edf74c..ecf450d8968 100644
--- a/e2e/components/BranchName.test.ts
+++ b/e2e/components/BranchName.test.ts
@@ -39,19 +39,5 @@ test.describe('BranchName', () => {
         })
       })
     }
-
-    test.describe('with forced underlines', () => {
-      test('default @vrt', async ({page}) => {
-        await visit(page, {
-          id: 'components-branchname--default',
-          globals: {
-            colorScheme: 'light',
-            prefersLinkUnderlines: 'true',
-          },
-        })
-
-        expect(await page.screenshot()).toMatchSnapshot('BranchName.Default.light.forcedUnderlines.png')
-      })
-    })
   })
 })
diff --git a/e2e/components/Breadcrumbs.test.ts b/e2e/components/Breadcrumbs.test.ts
index 17c4f1b4ea9..5b4a763084e 100644
--- a/e2e/components/Breadcrumbs.test.ts
+++ b/e2e/components/Breadcrumbs.test.ts
@@ -43,19 +43,5 @@ test.describe('Breadcrumbs', () => {
         })
       })
     }
-
-    test.describe('with forced underlines', () => {
-      test('default @vrt', async ({page}) => {
-        await visit(page, {
-          id: 'components-breadcrumbs--default',
-          globals: {
-            colorScheme: 'light',
-            prefersLinkUnderlines: 'true',
-          },
-        })
-
-        expect(await page.screenshot()).toMatchSnapshot('Breadcrumbs.Default.light.forcedUnderlines.png')
-      })
-    })
   })
 })
diff --git a/e2e/components/Link.test.ts b/e2e/components/Link.test.ts
index d556729b83f..75259a13e91 100644
--- a/e2e/components/Link.test.ts
+++ b/e2e/components/Link.test.ts
@@ -43,20 +43,6 @@ test.describe('Link', () => {
         })
       })
     }
-
-    test.describe('with forced underlines', () => {
-      test('default @vrt', async ({page}) => {
-        await visit(page, {
-          id: 'components-link--default',
-          globals: {
-            colorScheme: 'light',
-            prefersLinkUnderlines: 'true',
-          },
-        })
-
-        expect(await page.screenshot()).toMatchSnapshot('Link.Default.light.forcedUnderlines.png')
-      })
-    })
   })
 
   test.describe('Muted', () => {
@@ -99,20 +85,6 @@ test.describe('Link', () => {
         })
       })
     }
-
-    test.describe('with forced underlines', () => {
-      test('default @vrt', async ({page}) => {
-        await visit(page, {
-          id: 'components-link-features--muted',
-          globals: {
-            colorScheme: 'light',
-            prefersLinkUnderlines: 'true',
-          },
-        })
-
-        expect(await page.screenshot()).toMatchSnapshot('Link.Muted.light.forcedUnderlines.png')
-      })
-    })
   })
 
   test.describe('Underline', () => {
@@ -155,19 +127,5 @@ test.describe('Link', () => {
         })
       })
     }
-
-    test.describe('with forced underlines', () => {
-      test('default @vrt', async ({page}) => {
-        await visit(page, {
-          id: 'components-link-features--underline',
-          globals: {
-            colorScheme: 'light',
-            prefersLinkUnderlines: 'true',
-          },
-        })
-
-        expect(await page.screenshot()).toMatchSnapshot('Link.Underline.light.forcedUnderlines.png')
-      })
-    })
   })
 })
diff --git a/e2e/components/LinkButton.test.ts b/e2e/components/LinkButton.test.ts
index b9f46f25729..31e0c906172 100644
--- a/e2e/components/LinkButton.test.ts
+++ b/e2e/components/LinkButton.test.ts
@@ -137,20 +137,6 @@ test.describe('LinkButton', () => {
         })
       })
     }
-
-    test.describe('with forced underlines', () => {
-      test('default @vrt', async ({page}) => {
-        await visit(page, {
-          id: 'components-linkbutton-features--invisible',
-          globals: {
-            colorScheme: 'light',
-            prefersLinkUnderlines: 'true',
-          },
-        })
-
-        expect(await page.screenshot()).toMatchSnapshot('LinkButton.Invisible.light.forcedUnderlines.png')
-      })
-    })
   })
 
   test.describe('Large', () => {
diff --git a/e2e/test-helpers/storybook.ts b/e2e/test-helpers/storybook.ts
index 08eb3bafa4a..755e8848b12 100644
--- a/e2e/test-helpers/storybook.ts
+++ b/e2e/test-helpers/storybook.ts
@@ -28,7 +28,7 @@ export async function visit(page: Page, options: Options) {
     let params = ''
     for (const [key, value] of Object.entries(globals)) {
       if (params !== '') {
-        params += ';'
+        params += '&'
       }
       params += `${key}:${value}`
     }
diff --git a/src/ActionList/LinkItem.tsx b/src/ActionList/LinkItem.tsx
index d37c85ad270..d4decbe2292 100644
--- a/src/ActionList/LinkItem.tsx
+++ b/src/ActionList/LinkItem.tsx
@@ -32,7 +32,6 @@ export const LinkItem = React.forwardRef(({sx = {}, active, as: Component, ...pr
 
     // inherit Item styles
     color: 'inherit',
-    textDecoration: 'none',
     '&:hover': {color: 'inherit', textDecoration: 'none'},
   }
 
diff --git a/src/BaseStyles.tsx b/src/BaseStyles.tsx
index 896aef31b09..aece80f6087 100644
--- a/src/BaseStyles.tsx
+++ b/src/BaseStyles.tsx
@@ -26,13 +26,6 @@ const GlobalStyle = createGlobalStyle<{colorScheme?: 'light' | 'dark'}>`
   details-dialog:focus:not(:focus-visible):not(.focus-visible) {
     outline: none;
   }
-
-  /* Used to fake conditional styles using a technique by Lea Verou: https://lea.verou.me/blog/2020/10/the-var-space-hack-to-toggle-multiple-values-with-one-custom-property/ */
-  /* We have to use a zero-width space character (\u200B) as the value instead of a regular whitespace character because styled-components strips out properties that just have a whitespace value. */
-  :root {--prefers-link-underlines: \u200B;}
-  [data-a11y-link-underlines='true'] {
-    --prefers-link-underlines: initial;
-  }
 `
 
 const Base = styled.div<SystemTypographyProps & SystemCommonProps>`
diff --git a/src/BranchName/BranchName.tsx b/src/BranchName/BranchName.tsx
index c05c17da9ce..0ef498253b3 100644
--- a/src/BranchName/BranchName.tsx
+++ b/src/BranchName/BranchName.tsx
@@ -11,7 +11,7 @@ const BranchName = styled.a<SxProp>`
   color: ${get('colors.accent.fg')};
   background-color: ${get('colors.accent.subtle')};
   border-radius: ${get('radii.2')};
-  text-decoration: var(--prefers-link-underlines, underline);
+  text-decoration: none;
 
   ${sx};
 `
diff --git a/src/BranchName/__tests__/__snapshots__/BranchName.test.tsx.snap b/src/BranchName/__tests__/__snapshots__/BranchName.test.tsx.snap
index 10832885435..67046d28a4e 100644
--- a/src/BranchName/__tests__/__snapshots__/BranchName.test.tsx.snap
+++ b/src/BranchName/__tests__/__snapshots__/BranchName.test.tsx.snap
@@ -9,8 +9,8 @@ exports[`BranchName renders consistently 1`] = `
   color: #0969da;
   background-color: #ddf4ff;
   border-radius: 6px;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 <a
diff --git a/src/Breadcrumbs/Breadcrumbs.tsx b/src/Breadcrumbs/Breadcrumbs.tsx
index 19ab17c413d..70c33a25823 100644
--- a/src/Breadcrumbs/Breadcrumbs.tsx
+++ b/src/Breadcrumbs/Breadcrumbs.tsx
@@ -66,7 +66,7 @@ const BreadcrumbsItem = styled.a.attrs<StyledBreadcrumbsItemProps>(props => ({
   color: ${get('colors.accent.fg')};
   display: inline-block;
   font-size: ${get('fontSizes.1')};
-  text-decoration: var(--prefers-link-underlines, underline);
+  text-decoration: none;
   &:hover {
     text-decoration: underline;
   }
diff --git a/src/Breadcrumbs/__tests__/__snapshots__/BreadcrumbsItem.test.tsx.snap b/src/Breadcrumbs/__tests__/__snapshots__/BreadcrumbsItem.test.tsx.snap
index d0bc7896195..39ec4605f7d 100644
--- a/src/Breadcrumbs/__tests__/__snapshots__/BreadcrumbsItem.test.tsx.snap
+++ b/src/Breadcrumbs/__tests__/__snapshots__/BreadcrumbsItem.test.tsx.snap
@@ -5,8 +5,8 @@ exports[`Breadcrumbs.Item adds activeClassName={SELECTED_CLASS} when it gets a "
   color: #0969da;
   display: inline-block;
   font-size: 14px;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
@@ -32,8 +32,8 @@ exports[`Breadcrumbs.Item renders consistently 1`] = `
   color: #0969da;
   display: inline-block;
   font-size: 14px;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
@@ -57,8 +57,8 @@ exports[`Breadcrumbs.Item respects the "selected" prop 1`] = `
   color: #0969da;
   display: inline-block;
   font-size: 14px;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
diff --git a/src/Button/__tests__/__snapshots__/Button.test.tsx.snap b/src/Button/__tests__/__snapshots__/Button.test.tsx.snap
index 29133a6d684..bbaf6d6eba5 100644
--- a/src/Button/__tests__/__snapshots__/Button.test.tsx.snap
+++ b/src/Button/__tests__/__snapshots__/Button.test.tsx.snap
@@ -1512,13 +1512,11 @@ exports[`Button styles invisible button appropriately 1`] = `
   display: -webkit-inline-flex;
   display: -ms-inline-flexbox;
   display: inline-flex;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
 }
 
 .c0[href]:hover {
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
diff --git a/src/Button/styles.ts b/src/Button/styles.ts
index 9e4fdbce43e..426e5251d52 100644
--- a/src/Button/styles.ts
+++ b/src/Button/styles.ts
@@ -109,12 +109,6 @@ export const getVariantStyles = (variant: VariantType = 'default', theme?: Theme
       backgroundColor: 'transparent',
       borderColor: 'transparent',
       boxShadow: 'none',
-      '&[href]': {
-        textDecoration: 'var(--prefers-link-underlines, underline)',
-        '&:hover': {
-          textDecoration: 'var(--prefers-link-underlines, underline)',
-        },
-      },
       '&:hover:not([disabled])': {
         backgroundColor: 'btn.hoverBg',
       },
diff --git a/src/Heading/Heading.tsx b/src/Heading/Heading.tsx
index ce755302cd0..25a224aca34 100644
--- a/src/Heading/Heading.tsx
+++ b/src/Heading/Heading.tsx
@@ -39,7 +39,6 @@ const Heading = forwardRef(({as: Component = 'h2', ...props}, forwardedRef) => {
   return (
     <StyledHeading
       as={Component}
-      className="pc-Heading"
       {...props}
       // @ts-ignore shh
       ref={innerRef}
diff --git a/src/Heading/__tests__/__snapshots__/Heading.test.tsx.snap b/src/Heading/__tests__/__snapshots__/Heading.test.tsx.snap
index a2c8c37842d..0b34eccccc3 100644
--- a/src/Heading/__tests__/__snapshots__/Heading.test.tsx.snap
+++ b/src/Heading/__tests__/__snapshots__/Heading.test.tsx.snap
@@ -8,6 +8,6 @@ exports[`Heading renders consistently 1`] = `
 }
 
 <h2
-  className="c0 pc-Heading"
+  className="c0"
 />
 `;
diff --git a/src/Link/Link.tsx b/src/Link/Link.tsx
index fc6f3a73643..735c1674466 100644
--- a/src/Link/Link.tsx
+++ b/src/Link/Link.tsx
@@ -22,9 +22,9 @@ const hoverColor = system({
 
 const StyledLink = styled.a<StyledLinkProps>`
   color: ${props => (props.muted ? get('colors.fg.muted')(props) : get('colors.accent.fg')(props))};
-  text-decoration: ${props => (props.underline ? 'underline' : 'var(--prefers-link-underlines, underline)')};
+  text-decoration: ${props => (props.underline ? 'underline' : 'none')};
   &:hover {
-    text-decoration: ${props => (props.muted ? 'var(--prefers-link-underlines, underline)' : 'underline')};
+    text-decoration: ${props => (props.muted ? 'none' : 'underline')};
     ${props => (props.hoverColor ? hoverColor : props.muted ? `color: ${get('colors.accent.fg')(props)}` : '')};
   }
   &:is(button) {
diff --git a/src/Link/__tests__/__snapshots__/Link.test.tsx.snap b/src/Link/__tests__/__snapshots__/Link.test.tsx.snap
index 87a83747c45..eaad8694936 100644
--- a/src/Link/__tests__/__snapshots__/Link.test.tsx.snap
+++ b/src/Link/__tests__/__snapshots__/Link.test.tsx.snap
@@ -3,8 +3,8 @@
 exports[`Link applies button styles when rendering a button element 1`] = `
 .c0 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
@@ -37,8 +37,8 @@ exports[`Link applies button styles when rendering a button element 1`] = `
 exports[`Link passes href down to link element 1`] = `
 .c0 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
@@ -72,8 +72,8 @@ exports[`Link passes href down to link element 1`] = `
 exports[`Link renders consistently 1`] = `
 .c0 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
@@ -106,8 +106,8 @@ exports[`Link renders consistently 1`] = `
 exports[`Link respects hoverColor prop 1`] = `
 .c0 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
@@ -141,14 +141,14 @@ exports[`Link respects hoverColor prop 1`] = `
 exports[`Link respects the  "sx" prop when "muted" prop is also passed 1`] = `
 .c0 {
   color: #656d76;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   color: #ffffff;
 }
 
 .c0:hover {
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   color: #0969da;
 }
 
@@ -178,13 +178,13 @@ exports[`Link respects the  "sx" prop when "muted" prop is also passed 1`] = `
 exports[`Link respects the "muted" prop 1`] = `
 .c0 {
   color: #656d76;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   color: #0969da;
 }
 
diff --git a/src/NavList/__snapshots__/NavList.test.tsx.snap b/src/NavList/__snapshots__/NavList.test.tsx.snap
index cc1f8de9391..d5cd78ceef7 100644
--- a/src/NavList/__snapshots__/NavList.test.tsx.snap
+++ b/src/NavList/__snapshots__/NavList.test.tsx.snap
@@ -196,8 +196,8 @@ exports[`NavList renders a simple list 1`] = `
 
 .c3 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   padding-left: 8px;
   padding-right: 8px;
   padding-top: 6px;
@@ -212,8 +212,6 @@ exports[`NavList renders a simple list 1`] = `
   flex-grow: 1;
   border-radius: 6px;
   color: inherit;
-  -webkit-text-decoration: none;
-  text-decoration: none;
 }
 
 .c3:hover {
@@ -608,8 +606,8 @@ exports[`NavList renders with groups 1`] = `
 
 .c7 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   padding-left: 8px;
   padding-right: 8px;
   padding-top: 6px;
@@ -624,8 +622,6 @@ exports[`NavList renders with groups 1`] = `
   flex-grow: 1;
   border-radius: 6px;
   color: inherit;
-  -webkit-text-decoration: none;
-  text-decoration: none;
 }
 
 .c7:hover {
@@ -1095,8 +1091,8 @@ exports[`NavList.Item with NavList.SubNav does not have active styles if SubNav
 
 .c11 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   padding-left: 16px;
   padding-right: 8px;
   padding-top: 6px;
@@ -1111,8 +1107,6 @@ exports[`NavList.Item with NavList.SubNav does not have active styles if SubNav
   flex-grow: 1;
   border-radius: 6px;
   color: inherit;
-  -webkit-text-decoration: none;
-  text-decoration: none;
   font-size: 12px;
   font-weight: 400;
 }
@@ -1579,8 +1573,8 @@ exports[`NavList.Item with NavList.SubNav has active styles if SubNav contains t
 
 .c11 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   padding-left: 16px;
   padding-right: 8px;
   padding-top: 6px;
@@ -1595,8 +1589,6 @@ exports[`NavList.Item with NavList.SubNav has active styles if SubNav contains t
   flex-grow: 1;
   border-radius: 6px;
   color: inherit;
-  -webkit-text-decoration: none;
-  text-decoration: none;
   font-size: 12px;
   font-weight: 400;
 }
diff --git a/src/UnderlineNav2/__snapshots__/UnderlineNav.test.tsx.snap b/src/UnderlineNav2/__snapshots__/UnderlineNav.test.tsx.snap
index e2683254421..7087305ccb7 100644
--- a/src/UnderlineNav2/__snapshots__/UnderlineNav.test.tsx.snap
+++ b/src/UnderlineNav2/__snapshots__/UnderlineNav.test.tsx.snap
@@ -94,8 +94,8 @@ exports[`UnderlineNav renders consistently 1`] = `
 
 .c4 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   position: relative;
   display: -webkit-inline-box;
   display: -webkit-inline-flex;
@@ -182,8 +182,8 @@ exports[`UnderlineNav renders consistently 1`] = `
 
 .c7 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
   position: relative;
   display: -webkit-inline-box;
   display: -webkit-inline-flex;
diff --git a/src/__tests__/__snapshots__/SideNav.test.tsx.snap b/src/__tests__/__snapshots__/SideNav.test.tsx.snap
index 37336e1d5ec..d54bec24567 100644
--- a/src/__tests__/__snapshots__/SideNav.test.tsx.snap
+++ b/src/__tests__/__snapshots__/SideNav.test.tsx.snap
@@ -3,8 +3,8 @@
 exports[`SideNav SideNav.Link renders consistently 1`] = `
 .c0 {
   color: #0969da;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c0:hover {
diff --git a/src/__tests__/__snapshots__/TextInput.test.tsx.snap b/src/__tests__/__snapshots__/TextInput.test.tsx.snap
index 72d9a22ed98..8b9b35e4245 100644
--- a/src/__tests__/__snapshots__/TextInput.test.tsx.snap
+++ b/src/__tests__/__snapshots__/TextInput.test.tsx.snap
@@ -1710,13 +1710,11 @@ exports[`TextInput renders trailingAction icon button 1`] = `
   display: -webkit-inline-flex;
   display: -ms-inline-flexbox;
   display: inline-flex;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
 }
 
 .c4[href]:hover {
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c4:hover {
@@ -2361,13 +2359,11 @@ exports[`TextInput renders trailingAction text button 1`] = `
   display: -webkit-inline-flex;
   display: -ms-inline-flexbox;
   display: inline-flex;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
 }
 
 .c3[href]:hover {
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c3:hover {
@@ -2764,13 +2760,11 @@ exports[`TextInput renders trailingAction text button with a tooltip 1`] = `
   display: -webkit-inline-flex;
   display: -ms-inline-flexbox;
   display: inline-flex;
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
 }
 
 .c4[href]:hover {
-  -webkit-text-decoration: var(--prefers-link-underlines,underline);
-  text-decoration: var(--prefers-link-underlines,underline);
+  -webkit-text-decoration: none;
+  text-decoration: none;
 }
 
 .c4:hover {
diff --git a/src/drafts/Button2/styles.ts b/src/drafts/Button2/styles.ts
index 240a8c0044f..474a9e81991 100644
--- a/src/drafts/Button2/styles.ts
+++ b/src/drafts/Button2/styles.ts
@@ -106,12 +106,6 @@ export const getVariantStyles = (variant: VariantType = 'default', theme?: Theme
       backgroundColor: 'transparent',
       borderColor: 'transparent',
       boxShadow: 'none',
-      '&[href]': {
-        textDecoration: 'var(--prefers-link-underlines, underline)',
-        '&:hover': {
-          textDecoration: 'var(--prefers-link-underlines, underline)',
-        },
-      },
       '&:hover:not([disabled])': {
         backgroundColor: 'btn.hoverBg',
       },
diff --git a/src/utils/story-helpers.tsx b/src/utils/story-helpers.tsx
index 9a3719e242c..a48dcea9208 100644
--- a/src/utils/story-helpers.tsx
+++ b/src/utils/story-helpers.tsx
@@ -9,7 +9,7 @@ import {Icon} from '@primer/octicons-react'
 // we don't import StoryContext from storybook because of exports that conflict
 // with primer/react more: https://github.com/primer/react/runs/6129115026?check_suite_focus=true
 type StoryContext = Record<string, unknown> & {
-  globals: {colorScheme: string; showSurroundingElements?: boolean; prefersLinkUnderlines?: 'true' | 'false'}
+  globals: {colorScheme: string; showSurroundingElements?: boolean}
   parameters: Record<string, unknown>
 }
 
@@ -59,7 +59,7 @@ export const withThemeProvider = (Story: React.FC<React.PropsWithChildren<StoryC
   // used for testing ThemeProvider.stories.tsx
   if (context.parameters.disableThemeDecorator) return Story(context)
 
-  const {colorScheme, prefersLinkUnderlines} = context.globals
+  const {colorScheme} = context.globals
 
   if (colorScheme === 'all') {
     return (
@@ -87,9 +87,7 @@ export const withThemeProvider = (Story: React.FC<React.PropsWithChildren<StoryC
                   color: 'fg.default',
                 }}
               >
-                <div id={`html-addon-root-${scheme}`} data-a11y-link-underlines={prefersLinkUnderlines}>
-                  {Story(context)}
-                </div>
+                <div id={`html-addon-root-${scheme}`}>{Story(context)}</div>
               </Box>
             </BaseStyles>
           </ThemeProvider>
@@ -102,9 +100,7 @@ export const withThemeProvider = (Story: React.FC<React.PropsWithChildren<StoryC
     <ThemeProvider colorMode="day" dayScheme={colorScheme}>
       <GlobalStyle />
       <BaseStyles>
-        <div id="html-addon-root" data-a11y-link-underlines={prefersLinkUnderlines}>
-          {Story(context)}
-        </div>
+        <div id="html-addon-root">{Story(context)}</div>
       </BaseStyles>
     </ThemeProvider>
   )
@@ -122,16 +118,6 @@ export const toolbarTypes = {
     },
     showSurroundingElements: {},
   },
-  prefersLinkUnderlines: {
-    name: 'Force link underlines',
-    description: 'Whether to force underlines on links that are not otherwise underlined',
-    defaultValue: 'false',
-    toolbar: {
-      icon: 'link',
-      items: ['true', 'false'],
-      title: 'Force link underlines',
-    },
-  },
 }
 
 export const inputWrapperArgTypes: ArgTypes = {