diff --git a/Libraries/Components/View/ViewPropTypes.js b/Libraries/Components/View/ViewPropTypes.js
index c7ee1baa63a16b..5e558f9e17de39 100644
--- a/Libraries/Components/View/ViewPropTypes.js
+++ b/Libraries/Components/View/ViewPropTypes.js
@@ -461,6 +461,23 @@ export type ViewProps = $ReadOnly<{|
    */
   accessibilityActions?: ?$ReadOnlyArray<AccessibilityActionInfo>,
 
+  /**
+   *
+   * Node Information of a FlatList, VirtualizedList or SectionList collection item.
+   * A collection item starts at a given row and column in the collection, and spans one or more rows and columns.
+   *
+   * @platform android
+   *
+   */
+  accessibilityCollectionItem?: ?{
+    rowIndex: number,
+    rowSpan: number,
+    columnIndex: number,
+    columnSpan: number,
+    heading: boolean,
+    itemIndex: number,
+  },
+
   /**
    * Specifies the nativeID of the associated label text. When the assistive technology focuses on the component with this props, the text is read aloud.
    *
diff --git a/Libraries/Lists/FlatList.js b/Libraries/Lists/FlatList.js
index 2ebd0f673ab004..46f99c7603efa4 100644
--- a/Libraries/Lists/FlatList.js
+++ b/Libraries/Lists/FlatList.js
@@ -625,11 +625,18 @@ class FlatList<ItemT> extends React.PureComponent<Props<ItemT>, void> {
           return (
             <View style={StyleSheet.compose(styles.row, columnWrapperStyle)}>
               {item.map((it, kk) => {
+                const itemIndex = index * cols + kk;
+                const accessibilityCollectionItem = {
+                  ...info.accessibilityCollectionItem,
+                  columnIndex: itemIndex % cols,
+                  itemIndex: itemIndex,
+                };
                 const element = renderer({
                   // $FlowFixMe[incompatible-call]
                   item: it,
-                  index: index * cols + kk,
+                  index: itemIndex,
                   separators: info.separators,
+                  accessibilityCollectionItem,
                 });
                 return element != null ? (
                   <React.Fragment key={kk}>{element}</React.Fragment>
@@ -660,6 +667,7 @@ class FlatList<ItemT> extends React.PureComponent<Props<ItemT>, void> {
     return (
       <VirtualizedList
         {...restProps}
+        numColumns={numColumns}
         getItem={this._getItem}
         getItemCount={this._getItemCount}
         keyExtractor={this._keyExtractor}
diff --git a/Libraries/Lists/VirtualizedList.js b/Libraries/Lists/VirtualizedList.js
index 7755718a3792d6..07f705fdd28c7f 100644
--- a/Libraries/Lists/VirtualizedList.js
+++ b/Libraries/Lists/VirtualizedList.js
@@ -42,6 +42,7 @@ const ScrollView = require('../Components/ScrollView/ScrollView');
 const View = require('../Components/View/View');
 const Batchinator = require('../Interaction/Batchinator');
 const ReactNative = require('../Renderer/shims/ReactNative');
+const Platform = require('../Utilities/Platform');
 const flattenStyle = require('../StyleSheet/flattenStyle');
 const StyleSheet = require('../StyleSheet/StyleSheet');
 const infoLog = require('../Utilities/infoLog');
@@ -74,6 +75,11 @@ type State = {
  * Use the following helper functions for default values
  */
 
+// numColumnsOrDefault(this.props.numColumns)
+function numColumnsOrDefault(numColumns: ?number) {
+  return numColumns ?? 1;
+}
+
 // horizontalOrDefault(this.props.horizontal)
 function horizontalOrDefault(horizontal: ?boolean) {
   return horizontal ?? false;
@@ -1010,10 +1016,35 @@ class VirtualizedList extends React.PureComponent<Props, State> {
     );
   }
 
+  _getCellsInItemCount = (props: Props) => {
+    const {getCellsInItemCount, data} = props;
+    if (getCellsInItemCount) {
+      return getCellsInItemCount(data);
+    }
+    if (Array.isArray(data)) {
+      return data.length;
+    }
+    return 0;
+  };
+
   /* $FlowFixMe[missing-local-annot] The type annotation(s) required by Flow's
    * LTI update could not be added via codemod */
   _defaultRenderScrollComponent = props => {
+    const {getItemCount, data} = props;
     const onRefresh = props.onRefresh;
+    const numColumns = numColumnsOrDefault(props.numColumns);
+    const accessibilityRole = Platform.select({
+      android: numColumns > 1 ? 'grid' : 'list',
+    });
+    const rowCount = getItemCount(data);
+    const accessibilityCollection = {
+      // over-ride _getCellsInItemCount to handle Objects or other data formats
+      // see https://bit.ly/35RKX7H
+      itemCount: this._getCellsInItemCount(props),
+      rowCount,
+      columnCount: numColumns,
+      hierarchical: false,
+    };
     if (this._isNestedWithSameOrientation()) {
       // $FlowFixMe[prop-missing] - Typing ReactNativeComponent revealed errors
       return <View {...props} />;
@@ -1028,6 +1059,8 @@ class VirtualizedList extends React.PureComponent<Props, State> {
         // $FlowFixMe[prop-missing] Invalid prop usage
         <ScrollView
           {...props}
+          accessibilityRole={accessibilityRole}
+          accessibilityCollection={accessibilityCollection}
           refreshControl={
             props.refreshControl == null ? (
               <RefreshControl
@@ -1043,8 +1076,14 @@ class VirtualizedList extends React.PureComponent<Props, State> {
         />
       );
     } else {
-      // $FlowFixMe[prop-missing] Invalid prop usage
-      return <ScrollView {...props} />;
+      return (
+        // $FlowFixMe[prop-missing] Invalid prop usage
+        <ScrollView
+          {...props}
+          accessibilityRole={accessibilityRole}
+          accessibilityCollection={accessibilityCollection}
+        />
+      );
     }
   };
 
@@ -1821,10 +1860,19 @@ class CellRenderer extends React.Component<
     }
 
     if (renderItem) {
+      const accessibilityCollectionItem = {
+        itemIndex: index,
+        rowIndex: index,
+        rowSpan: 1,
+        columnIndex: 0,
+        columnSpan: 1,
+        heading: false,
+      };
       return renderItem({
         item,
         index,
         separators: this._separators,
+        accessibilityCollectionItem,
       });
     }
 
diff --git a/Libraries/Lists/VirtualizedListProps.js b/Libraries/Lists/VirtualizedListProps.js
index 7bf0cd0f539778..35db2c8bc61b1e 100644
--- a/Libraries/Lists/VirtualizedListProps.js
+++ b/Libraries/Lists/VirtualizedListProps.js
@@ -27,10 +27,20 @@ export type Separators = {
   ...
 };
 
+export type AccessibilityCollectionItem = {
+  itemIndex: number,
+  rowIndex: number,
+  rowSpan: number,
+  columnIndex: number,
+  columnSpan: number,
+  heading: boolean,
+};
+
 export type RenderItemProps<ItemT> = {
   item: ItemT,
   index: number,
   separators: Separators,
+  accessibilityCollectionItem: AccessibilityCollectionItem,
   ...
 };
 
@@ -49,9 +59,19 @@ type RequiredProps = {|
    */
   getItem: (data: any, index: number) => ?Item,
   /**
-   * Determines how many items are in the data blob.
+   * Determines how many items (rows) are in the data blob.
    */
   getItemCount: (data: any) => number,
+  /**
+   * Determines how many cells are in the data blob
+   * see https://bit.ly/35RKX7H
+   */
+  getCellsInItemCount?: (data: any) => number,
+  /**
+   * The number of columns used in FlatList.
+   * The default of 1 is used in other components to calculate the accessibilityCollection prop.
+   */
+  numColumns?: ?number,
 |};
 type OptionalProps = {|
   renderItem?: ?RenderItemType<Item>,
diff --git a/Libraries/Lists/VirtualizedList_EXPERIMENTAL.js b/Libraries/Lists/VirtualizedList_EXPERIMENTAL.js
index 3e2682f15a2a79..cb79424dfbecdb 100644
--- a/Libraries/Lists/VirtualizedList_EXPERIMENTAL.js
+++ b/Libraries/Lists/VirtualizedList_EXPERIMENTAL.js
@@ -49,6 +49,7 @@ const ScrollView = require('../Components/ScrollView/ScrollView');
 const View = require('../Components/View/View');
 const Batchinator = require('../Interaction/Batchinator');
 const ReactNative = require('../Renderer/shims/ReactNative');
+const Platform = require('../Utilities/Platform');
 const flattenStyle = require('../StyleSheet/flattenStyle');
 const StyleSheet = require('../StyleSheet/StyleSheet');
 const infoLog = require('../Utilities/infoLog');
@@ -81,6 +82,11 @@ type State = {
  * Use the following helper functions for default values
  */
 
+// numColumnsOrDefault(this.props.numColumns)
+function numColumnsOrDefault(numColumns: ?number) {
+  return numColumns ?? 1;
+}
+
 // horizontalOrDefault(this.props.horizontal)
 function horizontalOrDefault(horizontal: ?boolean) {
   return horizontal ?? false;
@@ -1205,10 +1211,35 @@ class VirtualizedList extends React.PureComponent<Props, State> {
     );
   }
 
+  _getCellsInItemCount = (props: Props) => {
+    const {getCellsInItemCount, data} = props;
+    if (getCellsInItemCount) {
+      return getCellsInItemCount(data);
+    }
+    if (Array.isArray(data)) {
+      return data.length;
+    }
+    return 0;
+  };
+
   /* $FlowFixMe[missing-local-annot] The type annotation(s) required by Flow's
    * LTI update could not be added via codemod */
   _defaultRenderScrollComponent = props => {
+    const {getItemCount, data} = props;
     const onRefresh = props.onRefresh;
+    const numColumns = numColumnsOrDefault(props.numColumns);
+    const accessibilityRole = Platform.select({
+      android: numColumns > 1 ? 'grid' : 'list',
+    });
+    const rowCount = getItemCount(data);
+    const accessibilityCollection = {
+      // over-ride _getCellsInItemCount to handle Objects or other data formats
+      // see https://bit.ly/35RKX7H
+      itemCount: this._getCellsInItemCount(props),
+      rowCount,
+      columnCount: numColumns,
+      hierarchical: false,
+    };
     if (this._isNestedWithSameOrientation()) {
       // $FlowFixMe[prop-missing] - Typing ReactNativeComponent revealed errors
       return <View {...props} />;
@@ -1223,6 +1254,8 @@ class VirtualizedList extends React.PureComponent<Props, State> {
         // $FlowFixMe[prop-missing] Invalid prop usage
         <ScrollView
           {...props}
+          accessibilityRole={accessibilityRole}
+          accessibilityCollection={accessibilityCollection}
           refreshControl={
             props.refreshControl == null ? (
               <RefreshControl
@@ -1238,8 +1271,14 @@ class VirtualizedList extends React.PureComponent<Props, State> {
         />
       );
     } else {
-      // $FlowFixMe[prop-missing] Invalid prop usage
-      return <ScrollView {...props} />;
+      return (
+        // $FlowFixMe[prop-missing] Invalid prop usage
+        <ScrollView
+          {...props}
+          accessibilityRole={accessibilityRole}
+          accessibilityCollection={accessibilityCollection}
+        />
+      );
     }
   };
 
@@ -2037,10 +2076,19 @@ class CellRenderer extends React.Component<
     }
 
     if (renderItem) {
+      const accessibilityCollectionItem = {
+        itemIndex: index,
+        rowIndex: index,
+        rowSpan: 1,
+        columnIndex: 0,
+        columnSpan: 1,
+        heading: false,
+      };
       return renderItem({
         item,
         index,
         separators: this._separators,
+        accessibilityCollectionItem,
       });
     }
 
diff --git a/Libraries/Lists/VirtualizedSectionList.js b/Libraries/Lists/VirtualizedSectionList.js
index aa18f60f05c9e4..ffeb45450f4262 100644
--- a/Libraries/Lists/VirtualizedSectionList.js
+++ b/Libraries/Lists/VirtualizedSectionList.js
@@ -9,7 +9,7 @@
  */
 
 import type {ViewToken} from './ViewabilityHelper';
-
+import type {AccessibilityCollectionItem} from './VirtualizedListProps';
 import {keyExtractor as defaultKeyExtractor} from './VirtualizeUtils';
 import invariant from 'invariant';
 import * as React from 'react';
@@ -342,7 +342,16 @@ class VirtualizedSectionList<
   _renderItem =
     (listItemCount: number) =>
     // eslint-disable-next-line react/no-unstable-nested-components
-    ({item, index}: {item: Item, index: number, ...}) => {
+    ({
+      item,
+      index,
+      accessibilityCollectionItem,
+    }: {
+      item: Item,
+      index: number,
+      accessibilityCollectionItem: AccessibilityCollectionItem,
+      ...
+    }) => {
       const info = this._subExtractor(index);
       if (!info) {
         return null;
@@ -371,6 +380,7 @@ class VirtualizedSectionList<
             LeadingSeparatorComponent={
               infoIndex === 0 ? this.props.SectionSeparatorComponent : undefined
             }
+            accessibilityCollectionItem={accessibilityCollectionItem}
             cellKey={info.key}
             index={infoIndex}
             item={item}
@@ -486,6 +496,7 @@ type ItemWithSeparatorProps = $ReadOnly<{|
   updatePropsFor: (prevCellKey: string, value: Object) => void,
   renderItem: Function,
   inverted: boolean,
+  accessibilityCollectionItem: AccessibilityCollectionItem,
 |}>;
 
 function ItemWithSeparator(props: ItemWithSeparatorProps): React.Node {
@@ -503,6 +514,7 @@ function ItemWithSeparator(props: ItemWithSeparatorProps): React.Node {
     index,
     section,
     inverted,
+    accessibilityCollectionItem,
   } = props;
 
   const [leadingSeparatorHiglighted, setLeadingSeparatorHighlighted] =
@@ -576,6 +588,7 @@ function ItemWithSeparator(props: ItemWithSeparatorProps): React.Node {
     index,
     section,
     separators,
+    accessibilityCollectionItem,
   });
   const leadingSeparator = LeadingSeparatorComponent != null && (
     <LeadingSeparatorComponent
diff --git a/Libraries/Lists/__tests__/VirtualizedList-test.js b/Libraries/Lists/__tests__/VirtualizedList-test.js
index 0928fd57c98c02..39ce49db2efd94 100644
--- a/Libraries/Lists/__tests__/VirtualizedList-test.js
+++ b/Libraries/Lists/__tests__/VirtualizedList-test.js
@@ -22,6 +22,7 @@ if (useExperimentalList) {
   VirtualizedListInjection.inject(VirtualizedList_EXPERIMENTAL);
 }
 
+const View = require('../../Components/View/View');
 const VirtualizedList = require('../VirtualizedList');
 
 describe('VirtualizedList', () => {
@@ -1542,6 +1543,38 @@ it('calls _onCellLayout properly', () => {
   expect(mock).not.toHaveBeenCalledWith(event, 'i3', 2);
 });
 
+it('renders list with custom column count', () => {
+  const numColumns = 3;
+  const component = ReactTestRenderer.create(
+    <VirtualizedList
+      data={[{key: 'i1'}, {key: 'i2'}, {key: 'i3'}, {key: 'i4'}, {key: 'i5'}]}
+      renderItem={({item}) => {
+        return (
+          <View key={item.key}>
+            {item.items.map((subitem, index) => (
+              <item value={subitem.key} key={index} />
+            ))}
+          </View>
+        );
+      }}
+      getItem={(data, index) => {
+        const ret = {key: index, items: []};
+        for (let i = 0; i < numColumns; i++) {
+          const item = data[index * numColumns + i];
+          if (item != null) {
+            ret.items.push(item);
+          }
+        }
+        return ret;
+      }}
+      numColumns={numColumns}
+      getItemCount={data => Math.ceil(data.length / numColumns)}
+      getCellsInItemCount={data => data.length}
+    />,
+  );
+  expect(component).toMatchSnapshot();
+});
+
 if (useExperimentalList) {
   describe('VirtualizedList (Experimental functionality)', () => {
     it('keeps viewport below last focused rendered', () => {
diff --git a/Libraries/Lists/__tests__/__snapshots__/FlatList-test.js.snap b/Libraries/Lists/__tests__/__snapshots__/FlatList-test.js.snap
index c8b33bb98df2de..062bf63589c965 100644
--- a/Libraries/Lists/__tests__/__snapshots__/FlatList-test.js.snap
+++ b/Libraries/Lists/__tests__/__snapshots__/FlatList-test.js.snap
@@ -6,6 +6,14 @@ exports[`FlatList renders all the bells and whistles 1`] = `
   ListEmptyComponent={[Function]}
   ListFooterComponent={[Function]}
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 2,
+      "hierarchical": false,
+      "itemCount": 5,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -29,6 +37,7 @@ exports[`FlatList renders all the bells and whistles 1`] = `
   getItemCount={[Function]}
   getItemLayout={[Function]}
   keyExtractor={[Function]}
+  numColumns={2}
   onContentSizeChange={[Function]}
   onLayout={[Function]}
   onMomentumScrollBegin={[Function]}
@@ -121,6 +130,14 @@ exports[`FlatList renders all the bells and whistles 1`] = `
 
 exports[`FlatList renders empty list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -144,6 +161,14 @@ exports[`FlatList renders empty list 1`] = `
 
 exports[`FlatList renders null list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   getItem={[Function]}
   getItemCount={[Function]}
   keyExtractor={[Function]}
@@ -166,6 +191,14 @@ exports[`FlatList renders null list 1`] = `
 
 exports[`FlatList renders simple list (multiple columns) 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 2,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 2,
+    }
+  }
   data={
     Array [
       Object {
@@ -182,6 +215,7 @@ exports[`FlatList renders simple list (multiple columns) 1`] = `
   getItem={[Function]}
   getItemCount={[Function]}
   keyExtractor={[Function]}
+  numColumns={2}
   onContentSizeChange={[Function]}
   onLayout={[Function]}
   onMomentumScrollBegin={[Function]}
@@ -237,6 +271,14 @@ exports[`FlatList renders simple list (multiple columns) 1`] = `
 
 exports[`FlatList renders simple list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -298,6 +340,14 @@ exports[`FlatList renders simple list 1`] = `
 exports[`FlatList renders simple list using ListItemComponent (multiple columns) 1`] = `
 <RCTScrollView
   ListItemComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 2,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 2,
+    }
+  }
   data={
     Array [
       Object {
@@ -314,6 +364,7 @@ exports[`FlatList renders simple list using ListItemComponent (multiple columns)
   getItem={[Function]}
   getItemCount={[Function]}
   keyExtractor={[Function]}
+  numColumns={2}
   onContentSizeChange={[Function]}
   onLayout={[Function]}
   onMomentumScrollBegin={[Function]}
@@ -369,6 +420,14 @@ exports[`FlatList renders simple list using ListItemComponent (multiple columns)
 exports[`FlatList renders simple list using ListItemComponent 1`] = `
 <RCTScrollView
   ListItemComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
diff --git a/Libraries/Lists/__tests__/__snapshots__/SectionList-test.js.snap b/Libraries/Lists/__tests__/__snapshots__/SectionList-test.js.snap
index 8e63455db938e0..139204914c5315 100644
--- a/Libraries/Lists/__tests__/__snapshots__/SectionList-test.js.snap
+++ b/Libraries/Lists/__tests__/__snapshots__/SectionList-test.js.snap
@@ -2,6 +2,14 @@
 
 exports[`SectionList rendering empty section headers is fine 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 4,
+    }
+  }
   data={
     Array [
       Object {
@@ -66,6 +74,14 @@ exports[`SectionList rendering empty section headers is fine 1`] = `
 
 exports[`SectionList renders a footer when there is no data 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 2,
+    }
+  }
   data={
     Array [
       Object {
@@ -115,6 +131,14 @@ exports[`SectionList renders a footer when there is no data 1`] = `
 
 exports[`SectionList renders a footer when there is no data and no header 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 2,
+    }
+  }
   data={
     Array [
       Object {
@@ -163,6 +187,14 @@ exports[`SectionList renders all the bells and whistles 1`] = `
   ListEmptyComponent={[Function]}
   ListFooterComponent={[Function]}
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 12,
+    }
+  }
   data={
     Array [
       Object {
@@ -258,7 +290,7 @@ exports[`SectionList renders all the bells and whistles 1`] = `
           v="highlighted:false,leadingItem:undefined,leadingSection:undefined,section:s1,trailingItem:i1s1,trailingSection:s2"
         />
         <itemForSection1
-          v="item:i1s1,index:0,section:s1,separators:[object Object]"
+          v="item:i1s1,index:0,section:s1,separators:[object Object],accessibilityCollectionItem:[object Object]"
         />
         <itemSeparatorForSection1
           v="highlighted:false,leadingItem:i1s1,leadingSection:undefined,section:s1,trailingItem:i2s1,trailingSection:s2"
@@ -271,7 +303,7 @@ exports[`SectionList renders all the bells and whistles 1`] = `
     >
       <View>
         <itemForSection1
-          v="item:i2s1,index:1,section:s1,separators:[object Object]"
+          v="item:i2s1,index:1,section:s1,separators:[object Object],accessibilityCollectionItem:[object Object]"
         />
         <sectionSeparator
           v="highlighted:false,leadingItem:i2s1,leadingSection:undefined,section:s1,trailingItem:undefined,trailingSection:s2"
@@ -303,7 +335,7 @@ exports[`SectionList renders all the bells and whistles 1`] = `
           v="highlighted:false,leadingItem:undefined,leadingSection:s1,section:s2,trailingItem:i1s2,trailingSection:s3"
         />
         <defaultItem
-          v="item:i1s2,index:0,section:s2,separators:[object Object]"
+          v="item:i1s2,index:0,section:s2,separators:[object Object],accessibilityCollectionItem:[object Object]"
         />
         <defaultItemSeparator
           v="highlighted:false,leadingItem:i1s2,leadingSection:s1,section:s2,trailingItem:i2s2,trailingSection:s3"
@@ -316,7 +348,7 @@ exports[`SectionList renders all the bells and whistles 1`] = `
     >
       <View>
         <defaultItem
-          v="item:i2s2,index:1,section:s2,separators:[object Object]"
+          v="item:i2s2,index:1,section:s2,separators:[object Object],accessibilityCollectionItem:[object Object]"
         />
         <sectionSeparator
           v="highlighted:false,leadingItem:i2s2,leadingSection:s1,section:s2,trailingItem:undefined,trailingSection:s3"
@@ -348,7 +380,7 @@ exports[`SectionList renders all the bells and whistles 1`] = `
           v="highlighted:false,leadingItem:undefined,leadingSection:s2,section:s3,trailingItem:i1s3,trailingSection:undefined"
         />
         <defaultItem
-          v="item:i1s3,index:0,section:s3,separators:[object Object]"
+          v="item:i1s3,index:0,section:s3,separators:[object Object],accessibilityCollectionItem:[object Object]"
         />
         <defaultItemSeparator
           v="highlighted:false,leadingItem:i1s3,leadingSection:s2,section:s3,trailingItem:i2s3,trailingSection:undefined"
@@ -361,7 +393,7 @@ exports[`SectionList renders all the bells and whistles 1`] = `
     >
       <View>
         <defaultItem
-          v="item:i2s3,index:1,section:s3,separators:[object Object]"
+          v="item:i2s3,index:1,section:s3,separators:[object Object],accessibilityCollectionItem:[object Object]"
         />
         <sectionSeparator
           v="highlighted:false,leadingItem:i2s3,leadingSection:s2,section:s3,trailingItem:undefined,trailingSection:undefined"
@@ -389,6 +421,14 @@ exports[`SectionList renders all the bells and whistles 1`] = `
 
 exports[`SectionList renders empty list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
diff --git a/Libraries/Lists/__tests__/__snapshots__/VirtualizedList-test.js.snap b/Libraries/Lists/__tests__/__snapshots__/VirtualizedList-test.js.snap
index c66d68042b8eab..997f255a1dfa6c 100644
--- a/Libraries/Lists/__tests__/__snapshots__/VirtualizedList-test.js.snap
+++ b/Libraries/Lists/__tests__/__snapshots__/VirtualizedList-test.js.snap
@@ -3,6 +3,14 @@
 exports[`VirtualizedList forwards correct stickyHeaderIndices when ListHeaderComponent present 1`] = `
 <RCTScrollView
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -149,6 +157,14 @@ exports[`VirtualizedList forwards correct stickyHeaderIndices when ListHeaderCom
 
 exports[`VirtualizedList forwards correct stickyHeaderIndices when all in initial render window 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -290,6 +306,14 @@ exports[`VirtualizedList forwards correct stickyHeaderIndices when all in initia
 
 exports[`VirtualizedList forwards correct stickyHeaderIndices when partially in initial render window 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -399,6 +423,14 @@ exports[`VirtualizedList forwards correct stickyHeaderIndices when partially in
 
 exports[`VirtualizedList handles nested lists 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 2,
+      "rowCount": 2,
+    }
+  }
   data={
     Array [
       Object {
@@ -475,6 +507,14 @@ exports[`VirtualizedList handles nested lists 1`] = `
       style={null}
     >
       <RCTScrollView
+        accessibilityCollection={
+          Object {
+            "columnCount": 1,
+            "hierarchical": false,
+            "itemCount": 2,
+            "rowCount": 2,
+          }
+        }
         data={
           Array [
             Object {
@@ -540,6 +580,14 @@ exports[`VirtualizedList handles nested lists 1`] = `
 exports[`VirtualizedList handles separators correctly 1`] = `
 <RCTScrollView
   ItemSeparatorComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -614,6 +662,14 @@ exports[`VirtualizedList handles separators correctly 1`] = `
 exports[`VirtualizedList handles separators correctly 2`] = `
 <RCTScrollView
   ItemSeparatorComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -688,6 +744,14 @@ exports[`VirtualizedList handles separators correctly 2`] = `
 exports[`VirtualizedList handles separators correctly 3`] = `
 <RCTScrollView
   ItemSeparatorComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -762,6 +826,14 @@ exports[`VirtualizedList handles separators correctly 3`] = `
 
 exports[`VirtualizedList keeps sticky headers above viewport visualized 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -980,6 +1052,14 @@ exports[`VirtualizedList renders all the bells and whistles 1`] = `
   ListEmptyComponent={[Function]}
   ListFooterComponent={[Function]}
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 5,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
@@ -1176,6 +1256,14 @@ exports[`VirtualizedList renders all the bells and whistles 1`] = `
 
 exports[`VirtualizedList renders empty list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -1196,6 +1284,14 @@ exports[`VirtualizedList renders empty list 1`] = `
 
 exports[`VirtualizedList renders empty list after batch 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -1219,6 +1315,14 @@ exports[`VirtualizedList renders empty list with empty component 1`] = `
   ListEmptyComponent={[Function]}
   ListFooterComponent={[Function]}
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -1252,6 +1356,14 @@ exports[`VirtualizedList renders empty list with empty component 1`] = `
 exports[`VirtualizedList renders list with empty component 1`] = `
 <RCTScrollView
   ListEmptyComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 1,
+    }
+  }
   data={
     Array [
       Object {
@@ -1287,6 +1399,14 @@ exports[`VirtualizedList renders list with empty component 1`] = `
 
 exports[`VirtualizedList renders null list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   getItem={[Function]}
   getItemCount={[Function]}
   onContentSizeChange={[Function]}
@@ -1306,6 +1426,14 @@ exports[`VirtualizedList renders null list 1`] = `
 
 exports[`VirtualizedList renders simple list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -1364,6 +1492,14 @@ exports[`VirtualizedList renders simple list 1`] = `
 exports[`VirtualizedList renders simple list using ListItemComponent 1`] = `
 <RCTScrollView
   ListItemComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -1420,6 +1556,14 @@ exports[`VirtualizedList renders simple list using ListItemComponent 1`] = `
 
 exports[`VirtualizedList renders sticky headers in viewport on batched render 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -1530,6 +1674,14 @@ exports[`VirtualizedList renders sticky headers in viewport on batched render 1`
 
 exports[`VirtualizedList test getItem functionality where data is not an Array 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 1,
+    }
+  }
   data={
     Map {
       "id_0" => Object {
@@ -1566,6 +1718,14 @@ exports[`VirtualizedList test getItem functionality where data is not an Array 1
 exports[`VirtualizedList warns if both renderItem or ListItemComponent are specified. Uses ListItemComponent 1`] = `
 <RCTScrollView
   ListItemComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 1,
+    }
+  }
   data={
     Array [
       Object {
@@ -1602,6 +1762,14 @@ exports[`VirtualizedList warns if both renderItem or ListItemComponent are speci
 
 exports[`adjusts render area with non-zero initialScrollIndex 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -1831,6 +1999,14 @@ exports[`adjusts render area with non-zero initialScrollIndex 1`] = `
 
 exports[`constrains batch render region when an item is removed 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 5,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
@@ -1908,6 +2084,14 @@ exports[`constrains batch render region when an item is removed 1`] = `
 
 exports[`discards intitial render if initialScrollIndex != 0 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -2080,6 +2264,14 @@ exports[`discards intitial render if initialScrollIndex != 0 1`] = `
 
 exports[`does not adjust render area until content area layed out 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -2209,6 +2401,14 @@ exports[`does not adjust render area until content area layed out 1`] = `
 
 exports[`does not over-render when there is less than initialNumToRender cells 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -2336,6 +2536,14 @@ exports[`does not over-render when there is less than initialNumToRender cells 1
 
 exports[`eventually renders all items when virtualization disabled 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -2466,6 +2674,14 @@ exports[`eventually renders all items when virtualization disabled 1`] = `
 
 exports[`expands first in viewport to render up to maxToRenderPerBatch on initial render 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -2573,6 +2789,14 @@ exports[`expands first in viewport to render up to maxToRenderPerBatch on initia
 
 exports[`expands render area by maxToRenderPerBatch on tick 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -2716,6 +2940,14 @@ exports[`expands render area by maxToRenderPerBatch on tick 1`] = `
 
 exports[`gracefully handles negaitve initialScrollIndex 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -2801,6 +3033,14 @@ exports[`gracefully handles negaitve initialScrollIndex 1`] = `
 
 exports[`initially renders nothing when initialNumToRender is 0 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -2871,6 +3111,14 @@ exports[`initially renders nothing when initialNumToRender is 0 1`] = `
 
 exports[`renders a zero-height tail spacer on initial render if getItemLayout not defined 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -2957,6 +3205,14 @@ exports[`renders a zero-height tail spacer on initial render if getItemLayout no
 
 exports[`renders full tail spacer if all cells measured 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3061,6 +3317,14 @@ exports[`renders full tail spacer if all cells measured 1`] = `
 
 exports[`renders initialNumToRender cells when virtualization disabled 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3161,6 +3425,14 @@ exports[`renders initialNumToRender cells when virtualization disabled 1`] = `
 
 exports[`renders items before initialScrollIndex on first batch tick when virtualization disabled 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3267,8 +3539,94 @@ exports[`renders items before initialScrollIndex on first batch tick when virtua
 </RCTScrollView>
 `;
 
+exports[`renders list with custom column count 1`] = `
+<RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 3,
+      "hierarchical": false,
+      "itemCount": 5,
+      "rowCount": 2,
+    }
+  }
+  data={
+    Array [
+      Object {
+        "key": "i1",
+      },
+      Object {
+        "key": "i2",
+      },
+      Object {
+        "key": "i3",
+      },
+      Object {
+        "key": "i4",
+      },
+      Object {
+        "key": "i5",
+      },
+    ]
+  }
+  getCellsInItemCount={[Function]}
+  getItem={[Function]}
+  getItemCount={[Function]}
+  numColumns={3}
+  onContentSizeChange={[Function]}
+  onLayout={[Function]}
+  onMomentumScrollBegin={[Function]}
+  onMomentumScrollEnd={[Function]}
+  onScroll={[Function]}
+  onScrollBeginDrag={[Function]}
+  onScrollEndDrag={[Function]}
+  renderItem={[Function]}
+  scrollEventThrottle={50}
+  stickyHeaderIndices={Array []}
+>
+  <View>
+    <View
+      onLayout={[Function]}
+      style={null}
+    >
+      <View>
+        <item
+          value="i1"
+        />
+        <item
+          value="i2"
+        />
+        <item
+          value="i3"
+        />
+      </View>
+    </View>
+    <View
+      onLayout={[Function]}
+      style={null}
+    >
+      <View>
+        <item
+          value="i4"
+        />
+        <item
+          value="i5"
+        />
+      </View>
+    </View>
+  </View>
+</RCTScrollView>
+`;
+
 exports[`renders new items when data is updated with non-zero initialScrollIndex 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 4,
+      "rowCount": 4,
+    }
+  }
   data={
     Array [
       Object {
@@ -3338,6 +3696,14 @@ exports[`renders new items when data is updated with non-zero initialScrollIndex
 
 exports[`renders no spacers up to initialScrollIndex on first render when virtualization disabled 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3411,6 +3777,14 @@ exports[`renders no spacers up to initialScrollIndex on first render when virtua
 
 exports[`renders offset cells in initial render when initialScrollIndex set 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3531,6 +3905,14 @@ exports[`renders offset cells in initial render when initialScrollIndex set 1`]
 
 exports[`renders tail spacer up to last measured index if getItemLayout not defined 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3635,6 +4017,14 @@ exports[`renders tail spacer up to last measured index if getItemLayout not defi
 
 exports[`renders tail spacer up to last measured with irregular layout when getItemLayout undefined 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3731,6 +4121,14 @@ exports[`renders tail spacer up to last measured with irregular layout when getI
 
 exports[`renders windowSize derived region at bottom 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3838,6 +4236,14 @@ exports[`renders windowSize derived region at bottom 1`] = `
 
 exports[`renders windowSize derived region at top 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3931,6 +4337,14 @@ exports[`renders windowSize derived region at top 1`] = `
 
 exports[`renders windowSize derived region in middle 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4059,6 +4473,14 @@ exports[`renders windowSize derived region in middle 1`] = `
 
 exports[`renders zero-height tail spacer on batch render if cells not yet measured and getItemLayout not defined 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4147,6 +4569,14 @@ exports[`renders zero-height tail spacer on batch render if cells not yet measur
 
 exports[`retains batch render region when an item is appended 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 11,
+      "rowCount": 11,
+    }
+  }
   data={
     Array [
       Object {
@@ -4284,6 +4714,14 @@ exports[`retains batch render region when an item is appended 1`] = `
 
 exports[`retains initial render region when an item is appended 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 11,
+      "rowCount": 11,
+    }
+  }
   data={
     Array [
       Object {
@@ -4371,6 +4809,14 @@ exports[`retains initial render region when an item is appended 1`] = `
 
 exports[`retains intitial render if initialScrollIndex == 0 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -4578,6 +5024,14 @@ exports[`retains intitial render if initialScrollIndex == 0 1`] = `
 
 exports[`unmounts sticky headers moved below viewport 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
diff --git a/Libraries/Lists/__tests__/__snapshots__/VirtualizedList_EXPERIMENTAL-test.js.snap b/Libraries/Lists/__tests__/__snapshots__/VirtualizedList_EXPERIMENTAL-test.js.snap
index 61cb26060262f6..9675a4e5bc0337 100644
--- a/Libraries/Lists/__tests__/__snapshots__/VirtualizedList_EXPERIMENTAL-test.js.snap
+++ b/Libraries/Lists/__tests__/__snapshots__/VirtualizedList_EXPERIMENTAL-test.js.snap
@@ -2,6 +2,14 @@
 
 exports[`VirtualizedList (Experimental functionality) keeps viewport above last focused rendered 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -200,6 +208,14 @@ exports[`VirtualizedList (Experimental functionality) keeps viewport above last
 
 exports[`VirtualizedList (Experimental functionality) keeps viewport below last focused rendered 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -414,6 +430,14 @@ exports[`VirtualizedList (Experimental functionality) keeps viewport below last
 
 exports[`VirtualizedList (Experimental functionality) virtualizes away last focused index if item removed 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 19,
+      "rowCount": 19,
+    }
+  }
   data={
     Array [
       Object {
@@ -601,6 +625,14 @@ exports[`VirtualizedList (Experimental functionality) virtualizes away last focu
 
 exports[`VirtualizedList (Experimental functionality) virtualizes away last focused item if focus changes to a new cell 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -768,6 +800,14 @@ exports[`VirtualizedList (Experimental functionality) virtualizes away last focu
 exports[`VirtualizedList forwards correct stickyHeaderIndices when ListHeaderComponent present 1`] = `
 <RCTScrollView
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -924,6 +964,14 @@ exports[`VirtualizedList forwards correct stickyHeaderIndices when ListHeaderCom
 
 exports[`VirtualizedList forwards correct stickyHeaderIndices when all in initial render window 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -1075,6 +1123,14 @@ exports[`VirtualizedList forwards correct stickyHeaderIndices when all in initia
 
 exports[`VirtualizedList forwards correct stickyHeaderIndices when partially in initial render window 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -1189,6 +1245,14 @@ exports[`VirtualizedList forwards correct stickyHeaderIndices when partially in
 
 exports[`VirtualizedList handles nested lists 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 2,
+      "rowCount": 2,
+    }
+  }
   data={
     Array [
       Object {
@@ -1269,6 +1333,14 @@ exports[`VirtualizedList handles nested lists 1`] = `
       style={null}
     >
       <RCTScrollView
+        accessibilityCollection={
+          Object {
+            "columnCount": 1,
+            "hierarchical": false,
+            "itemCount": 2,
+            "rowCount": 2,
+          }
+        }
         data={
           Array [
             Object {
@@ -1336,6 +1408,14 @@ exports[`VirtualizedList handles nested lists 1`] = `
 exports[`VirtualizedList handles separators correctly 1`] = `
 <RCTScrollView
   ItemSeparatorComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -1413,6 +1493,14 @@ exports[`VirtualizedList handles separators correctly 1`] = `
 exports[`VirtualizedList handles separators correctly 2`] = `
 <RCTScrollView
   ItemSeparatorComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -1490,6 +1578,14 @@ exports[`VirtualizedList handles separators correctly 2`] = `
 exports[`VirtualizedList handles separators correctly 3`] = `
 <RCTScrollView
   ItemSeparatorComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -1567,6 +1663,14 @@ exports[`VirtualizedList handles separators correctly 3`] = `
 
 exports[`VirtualizedList keeps sticky headers above viewport visualized 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -1754,6 +1858,14 @@ exports[`VirtualizedList renders all the bells and whistles 1`] = `
   ListEmptyComponent={[Function]}
   ListFooterComponent={[Function]}
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 5,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
@@ -1955,6 +2067,14 @@ exports[`VirtualizedList renders all the bells and whistles 1`] = `
 
 exports[`VirtualizedList renders empty list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -1975,6 +2095,14 @@ exports[`VirtualizedList renders empty list 1`] = `
 
 exports[`VirtualizedList renders empty list after batch 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -1998,6 +2126,14 @@ exports[`VirtualizedList renders empty list with empty component 1`] = `
   ListEmptyComponent={[Function]}
   ListFooterComponent={[Function]}
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -2031,6 +2167,14 @@ exports[`VirtualizedList renders empty list with empty component 1`] = `
 exports[`VirtualizedList renders list with empty component 1`] = `
 <RCTScrollView
   ListEmptyComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 1,
+    }
+  }
   data={
     Array [
       Object {
@@ -2067,6 +2211,14 @@ exports[`VirtualizedList renders list with empty component 1`] = `
 
 exports[`VirtualizedList renders null list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   getItem={[Function]}
   getItemCount={[Function]}
   onContentSizeChange={[Function]}
@@ -2086,6 +2238,14 @@ exports[`VirtualizedList renders null list 1`] = `
 
 exports[`VirtualizedList renders simple list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -2147,6 +2307,14 @@ exports[`VirtualizedList renders simple list 1`] = `
 exports[`VirtualizedList renders simple list using ListItemComponent 1`] = `
 <RCTScrollView
   ListItemComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 3,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -2206,6 +2374,14 @@ exports[`VirtualizedList renders simple list using ListItemComponent 1`] = `
 
 exports[`VirtualizedList renders sticky headers in viewport on batched render 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -2321,6 +2497,14 @@ exports[`VirtualizedList renders sticky headers in viewport on batched render 1`
 
 exports[`VirtualizedList test getItem functionality where data is not an Array 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 1,
+    }
+  }
   data={
     Map {
       "id_0" => Object {
@@ -2358,6 +2542,14 @@ exports[`VirtualizedList test getItem functionality where data is not an Array 1
 exports[`VirtualizedList warns if both renderItem or ListItemComponent are specified. Uses ListItemComponent 1`] = `
 <RCTScrollView
   ListItemComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 1,
+    }
+  }
   data={
     Array [
       Object {
@@ -2395,6 +2587,14 @@ exports[`VirtualizedList warns if both renderItem or ListItemComponent are speci
 
 exports[`adjusts render area with non-zero initialScrollIndex 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -2644,6 +2844,14 @@ exports[`adjusts render area with non-zero initialScrollIndex 1`] = `
 
 exports[`constrains batch render region when an item is removed 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 5,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
@@ -2726,6 +2934,14 @@ exports[`constrains batch render region when an item is removed 1`] = `
 
 exports[`discards intitial render if initialScrollIndex != 0 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -2869,6 +3085,14 @@ exports[`discards intitial render if initialScrollIndex != 0 1`] = `
 
 exports[`does not adjust render area until content area layed out 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -3003,6 +3227,14 @@ exports[`does not adjust render area until content area layed out 1`] = `
 
 exports[`does not over-render when there is less than initialNumToRender cells 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3115,6 +3347,14 @@ exports[`does not over-render when there is less than initialNumToRender cells 1
 
 exports[`eventually renders all items when virtualization disabled 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3255,6 +3495,14 @@ exports[`eventually renders all items when virtualization disabled 1`] = `
 
 exports[`expands first in viewport to render up to maxToRenderPerBatch on initial render 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3343,6 +3591,14 @@ exports[`expands first in viewport to render up to maxToRenderPerBatch on initia
 
 exports[`expands render area by maxToRenderPerBatch on tick 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -3493,6 +3749,14 @@ exports[`expands render area by maxToRenderPerBatch on tick 1`] = `
 
 exports[`gracefully handles negaitve initialScrollIndex 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3589,6 +3853,14 @@ exports[`gracefully handles negaitve initialScrollIndex 1`] = `
 
 exports[`initially renders nothing when initialNumToRender is 0 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3652,6 +3924,14 @@ exports[`initially renders nothing when initialNumToRender is 0 1`] = `
 
 exports[`renders a zero-height tail spacer on initial render if getItemLayout not defined 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3741,6 +4021,14 @@ exports[`renders a zero-height tail spacer on initial render if getItemLayout no
 
 exports[`renders full tail spacer if all cells measured 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3850,6 +4138,14 @@ exports[`renders full tail spacer if all cells measured 1`] = `
 
 exports[`renders initialNumToRender cells when virtualization disabled 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -3948,6 +4244,14 @@ exports[`renders initialNumToRender cells when virtualization disabled 1`] = `
 
 exports[`renders items before initialScrollIndex on first batch tick when virtualization disabled 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4061,8 +4365,96 @@ exports[`renders items before initialScrollIndex on first batch tick when virtua
 </RCTScrollView>
 `;
 
+exports[`renders list with custom column count 1`] = `
+<RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 3,
+      "hierarchical": false,
+      "itemCount": 5,
+      "rowCount": 2,
+    }
+  }
+  data={
+    Array [
+      Object {
+        "key": "i1",
+      },
+      Object {
+        "key": "i2",
+      },
+      Object {
+        "key": "i3",
+      },
+      Object {
+        "key": "i4",
+      },
+      Object {
+        "key": "i5",
+      },
+    ]
+  }
+  getCellsInItemCount={[Function]}
+  getItem={[Function]}
+  getItemCount={[Function]}
+  numColumns={3}
+  onContentSizeChange={[Function]}
+  onLayout={[Function]}
+  onMomentumScrollBegin={[Function]}
+  onMomentumScrollEnd={[Function]}
+  onScroll={[Function]}
+  onScrollBeginDrag={[Function]}
+  onScrollEndDrag={[Function]}
+  renderItem={[Function]}
+  scrollEventThrottle={50}
+  stickyHeaderIndices={Array []}
+>
+  <View>
+    <View
+      onFocusCapture={[Function]}
+      onLayout={[Function]}
+      style={null}
+    >
+      <View>
+        <item
+          value="i1"
+        />
+        <item
+          value="i2"
+        />
+        <item
+          value="i3"
+        />
+      </View>
+    </View>
+    <View
+      onFocusCapture={[Function]}
+      onLayout={[Function]}
+      style={null}
+    >
+      <View>
+        <item
+          value="i4"
+        />
+        <item
+          value="i5"
+        />
+      </View>
+    </View>
+  </View>
+</RCTScrollView>
+`;
+
 exports[`renders new items when data is updated with non-zero initialScrollIndex 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 4,
+      "rowCount": 4,
+    }
+  }
   data={
     Array [
       Object {
@@ -4136,6 +4528,14 @@ exports[`renders new items when data is updated with non-zero initialScrollIndex
 
 exports[`renders no spacers up to initialScrollIndex on first render when virtualization disabled 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4211,6 +4611,14 @@ exports[`renders no spacers up to initialScrollIndex on first render when virtua
 
 exports[`renders offset cells in initial render when initialScrollIndex set 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4314,6 +4722,14 @@ exports[`renders offset cells in initial render when initialScrollIndex set 1`]
 
 exports[`renders tail spacer up to last measured index if getItemLayout not defined 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4423,6 +4839,14 @@ exports[`renders tail spacer up to last measured index if getItemLayout not defi
 
 exports[`renders tail spacer up to last measured with irregular layout when getItemLayout undefined 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4523,6 +4947,14 @@ exports[`renders tail spacer up to last measured with irregular layout when getI
 
 exports[`renders windowSize derived region at bottom 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4636,6 +5068,14 @@ exports[`renders windowSize derived region at bottom 1`] = `
 
 exports[`renders windowSize derived region at top 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4733,6 +5173,14 @@ exports[`renders windowSize derived region at top 1`] = `
 
 exports[`renders windowSize derived region in middle 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4869,6 +5317,14 @@ exports[`renders windowSize derived region in middle 1`] = `
 
 exports[`renders zero-height tail spacer on batch render if cells not yet measured and getItemLayout not defined 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 10,
+      "rowCount": 10,
+    }
+  }
   data={
     Array [
       Object {
@@ -4960,6 +5416,14 @@ exports[`renders zero-height tail spacer on batch render if cells not yet measur
 
 exports[`retains batch render region when an item is appended 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 11,
+      "rowCount": 11,
+    }
+  }
   data={
     Array [
       Object {
@@ -5107,6 +5571,14 @@ exports[`retains batch render region when an item is appended 1`] = `
 
 exports[`retains initial render region when an item is appended 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 11,
+      "rowCount": 11,
+    }
+  }
   data={
     Array [
       Object {
@@ -5197,6 +5669,14 @@ exports[`retains initial render region when an item is appended 1`] = `
 
 exports[`retains intitial render if initialScrollIndex == 0 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
@@ -5380,6 +5860,14 @@ exports[`retains intitial render if initialScrollIndex == 0 1`] = `
 
 exports[`unmounts sticky headers moved below viewport 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 20,
+      "rowCount": 20,
+    }
+  }
   data={
     Array [
       Object {
diff --git a/Libraries/Lists/__tests__/__snapshots__/VirtualizedSectionList-test.js.snap b/Libraries/Lists/__tests__/__snapshots__/VirtualizedSectionList-test.js.snap
index d0fc9fa107c237..5c32fe8f76d8bb 100644
--- a/Libraries/Lists/__tests__/__snapshots__/VirtualizedSectionList-test.js.snap
+++ b/Libraries/Lists/__tests__/__snapshots__/VirtualizedSectionList-test.js.snap
@@ -2,6 +2,14 @@
 
 exports[`VirtualizedSectionList handles nested lists 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 4,
+    }
+  }
   data={
     Array [
       Object {
@@ -102,6 +110,14 @@ exports[`VirtualizedSectionList handles nested lists 1`] = `
       style={null}
     >
       <RCTScrollView
+        accessibilityCollection={
+          Object {
+            "columnCount": 1,
+            "hierarchical": false,
+            "itemCount": 1,
+            "rowCount": 4,
+          }
+        }
         data={
           Array [
             Object {
@@ -198,6 +214,14 @@ exports[`VirtualizedSectionList handles nested lists 1`] = `
 
 exports[`VirtualizedSectionList handles separators correctly 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
@@ -331,6 +355,14 @@ exports[`VirtualizedSectionList handles separators correctly 1`] = `
 
 exports[`VirtualizedSectionList handles separators correctly 2`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
@@ -464,6 +496,14 @@ exports[`VirtualizedSectionList handles separators correctly 2`] = `
 
 exports[`VirtualizedSectionList handles separators correctly 3`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
@@ -598,6 +638,14 @@ exports[`VirtualizedSectionList handles separators correctly 3`] = `
 
 exports[`VirtualizedSectionList handles separators correctly 4`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
@@ -735,6 +783,14 @@ exports[`VirtualizedSectionList renders all the bells and whistles 1`] = `
   ListEmptyComponent={[Function]}
   ListFooterComponent={[Function]}
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 7,
+    }
+  }
   data={
     Array [
       Object {
@@ -976,6 +1032,14 @@ exports[`VirtualizedSectionList renders all the bells and whistles 1`] = `
 
 exports[`VirtualizedSectionList renders empty list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -1000,6 +1064,14 @@ exports[`VirtualizedSectionList renders empty list with empty component 1`] = `
   ListEmptyComponent={[Function]}
   ListFooterComponent={[Function]}
   ListHeaderComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 0,
+      "rowCount": 0,
+    }
+  }
   data={Array []}
   getItem={[Function]}
   getItemCount={[Function]}
@@ -1034,6 +1106,14 @@ exports[`VirtualizedSectionList renders empty list with empty component 1`] = `
 exports[`VirtualizedSectionList renders list with empty component 1`] = `
 <RCTScrollView
   ListEmptyComponent={[Function]}
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 3,
+    }
+  }
   data={
     Array [
       Object {
@@ -1083,6 +1163,14 @@ exports[`VirtualizedSectionList renders list with empty component 1`] = `
 
 exports[`VirtualizedSectionList renders simple list 1`] = `
 <RCTScrollView
+  accessibilityCollection={
+    Object {
+      "columnCount": 1,
+      "hierarchical": false,
+      "itemCount": 1,
+      "rowCount": 5,
+    }
+  }
   data={
     Array [
       Object {
diff --git a/packages/rn-tester/js/examples/FlatList/FlatList-basic.js b/packages/rn-tester/js/examples/FlatList/FlatList-basic.js
index 9d3796062a7dff..704be0ae1c27a1 100644
--- a/packages/rn-tester/js/examples/FlatList/FlatList-basic.js
+++ b/packages/rn-tester/js/examples/FlatList/FlatList-basic.js
@@ -301,17 +301,21 @@ class FlatListExample extends React.PureComponent<Props, State> {
        * To see the error, delete this comment and run Flow. */
       /* $FlowFixMe[missing-local-annot] The type annotation(s) required by
        * Flow's LTI update could not be added via codemod */
-      [flatListPropKey]: ({item, separators}) => {
+      [flatListPropKey]: ({item, separators, accessibilityCollectionItem}) => {
         return (
-          <ItemComponent
-            item={item}
-            horizontal={this.state.horizontal}
-            fixedHeight={this.state.fixedHeight}
-            onPress={this._onPressCallback()}
-            onShowUnderlay={separators.highlight}
-            onHideUnderlay={separators.unhighlight}
-            textSelectable={this.state.textSelectable}
-          />
+          <View
+            importantForAccessibility="yes"
+            accessibilityCollectionItem={accessibilityCollectionItem}>
+            <ItemComponent
+              item={item}
+              horizontal={this.state.horizontal}
+              fixedHeight={this.state.fixedHeight}
+              onPress={this._onPressCallback()}
+              onShowUnderlay={separators.highlight}
+              onHideUnderlay={separators.unhighlight}
+              textSelectable={this.state.textSelectable}
+            />
+          </View>
         );
       },
     };
diff --git a/packages/rn-tester/js/examples/FlatList/FlatList-multiColumn.js b/packages/rn-tester/js/examples/FlatList/FlatList-multiColumn.js
index 66f0fc441532f5..e0bf2fb4e5eeed 100644
--- a/packages/rn-tester/js/examples/FlatList/FlatList-multiColumn.js
+++ b/packages/rn-tester/js/examples/FlatList/FlatList-multiColumn.js
@@ -10,7 +10,10 @@
 
 'use strict';
 
-import type {RenderItemProps} from 'react-native/Libraries/Lists/VirtualizedList';
+import type {
+  RenderItemProps,
+  AccessibilityCollectionItem,
+} from 'react-native/Libraries/Lists/VirtualizedListProps';
 import type {RNTesterModuleExample} from '../../types/RNTesterTypes';
 const RNTesterPage = require('../../components/RNTesterPage');
 const React = require('react');
@@ -142,9 +145,19 @@ class MultiColumnExample extends React.PureComponent<
       getItemLayout(data, index).length + 2 * (CARD_MARGIN + BORDER_WIDTH);
     return {length, offset: length * index, index};
   }
-  _renderItemComponent = ({item}: RenderItemProps<any | Item>) => {
+  _renderItemComponent = ({
+    item,
+    accessibilityCollectionItem,
+  }: {
+    item: Item,
+    accessibilityCollectionItem: AccessibilityCollectionItem,
+    ...
+  }) => {
     return (
-      <View style={styles.card}>
+      <View
+        importantForAccessibility="yes"
+        accessibilityCollectionItem={accessibilityCollectionItem}
+        style={styles.card}>
         <ItemComponent
           item={item}
           fixedHeight={this.state.fixedHeight}
diff --git a/packages/rn-tester/js/examples/FlatList/FlatList-nested.js b/packages/rn-tester/js/examples/FlatList/FlatList-nested.js
new file mode 100644
index 00000000000000..8d7d6f6de12441
--- /dev/null
+++ b/packages/rn-tester/js/examples/FlatList/FlatList-nested.js
@@ -0,0 +1,112 @@
+/**
+ * Copyright (c) Meta Platforms, Inc. and affiliates.
+ *
+ * This source code is licensed under the MIT license found in the
+ * LICENSE file in the root directory of this source tree.
+ *
+ * @format
+ * @flow
+ */
+
+import * as React from 'react';
+import {
+  SafeAreaView,
+  View,
+  FlatList,
+  StyleSheet,
+  Text,
+  StatusBar,
+} from 'react-native';
+
+import type {
+  RenderItemProps,
+  AccessibilityCollectionItem,
+} from 'react-native/Libraries/Lists/VirtualizedListProps';
+
+const DATA = [
+  'First Item',
+  'Second Item',
+  'Third Item',
+  'Fourth Item',
+  'Fifth Item',
+  'Sixth Item',
+  'Seven Item',
+  'Eight Item',
+  'Nine Item',
+  'Ten Item',
+];
+
+const Item = ({
+  item,
+  accessibilityCollectionItem,
+}: {
+  item: string,
+  accessibilityCollectionItem: AccessibilityCollectionItem,
+  ...
+}) => (
+  <View
+    importantForAccessibility="yes"
+    accessibilityCollectionItem={accessibilityCollectionItem}
+    style={styles.item}>
+    <Text style={styles.title}>{item}</Text>
+  </View>
+);
+
+const renderItem = ({
+  item,
+  separators,
+  accessibilityCollectionItem,
+}: RenderItemProps<string>) => (
+  <Item item={item} accessibilityCollectionItem={accessibilityCollectionItem} />
+);
+
+const renderFlatList = ({item}: RenderItemProps<number>) => {
+  return (
+    <View>
+      <Text>Flatlist {item}</Text>
+      <FlatList renderItem={renderItem} horizontal data={DATA} />
+    </View>
+  );
+};
+
+const FlatListNested = (): React.Node => {
+  return (
+    <SafeAreaView style={styles.container}>
+      <FlatList
+        data={[1, 2, 3]}
+        renderItem={renderFlatList}
+        keyExtractor={item => item.toString()}
+      />
+    </SafeAreaView>
+  );
+};
+
+const styles = StyleSheet.create({
+  container: {
+    flex: 1,
+    marginTop: StatusBar.currentHeight || 0,
+  },
+  item: {
+    backgroundColor: '#f9c2ff',
+    padding: 20,
+    marginVertical: 8,
+    marginHorizontal: 16,
+  },
+  title: {
+    fontSize: 16,
+  },
+});
+
+exports.title = 'FlatList Nested';
+exports.testTitle = 'Test accessibility announcement in nested flatlist';
+exports.category = 'ListView';
+exports.documentationURL = 'https://reactnative.dev/docs/flatlist';
+exports.description = 'Nested flatlist example';
+exports.examples = [
+  {
+    title: 'FlatList Nested example',
+    render: function (): React.Element<typeof FlatListNested> {
+      return <FlatListNested />;
+    },
+  },
+];
diff --git a/packages/rn-tester/js/examples/SectionList/SectionList-scrollable.js b/packages/rn-tester/js/examples/SectionList/SectionList-scrollable.js
index aa8ea52ec9899e..0398496a5f7596 100644
--- a/packages/rn-tester/js/examples/SectionList/SectionList-scrollable.js
+++ b/packages/rn-tester/js/examples/SectionList/SectionList-scrollable.js
@@ -9,6 +9,7 @@
  */
 
 'use strict';
+import type {AccessibilityCollectionItem} from 'react-native/Libraries/Lists/VirtualizedListProps';
 import type {Item} from '../../components/ListExampleShared';
 const RNTesterPage = require('../../components/RNTesterPage');
 const React = require('react');
@@ -118,7 +119,7 @@ const renderItemComponent =
   (setItemState: (item: Item) => void) =>
   /* $FlowFixMe[missing-local-annot] The type annotation(s) required by Flow's
    * LTI update could not be added via codemod */
-  ({item, separators}) => {
+  ({item, separators, accessibilityCollectionItem}) => {
     if (isNaN(item.key)) {
       return;
     }
@@ -128,12 +129,16 @@ const renderItemComponent =
     };
 
     return (
-      <ItemComponent
-        item={item}
-        onPress={onPress}
-        onHideUnderlay={separators.unhighlight}
-        onShowUnderlay={separators.highlight}
-      />
+      <View
+        importantForAccessibility="yes"
+        accessibilityCollectionItem={accessibilityCollectionItem}>
+        <ItemComponent
+          item={item}
+          onPress={onPress}
+          onHideUnderlay={separators.unhighlight}
+          onShowUnderlay={separators.highlight}
+        />
+      </View>
     );
   };
 
diff --git a/packages/rn-tester/js/utils/RNTesterList.android.js b/packages/rn-tester/js/utils/RNTesterList.android.js
index 97ffe9759a1b7c..f781797fe4a73f 100644
--- a/packages/rn-tester/js/utils/RNTesterList.android.js
+++ b/packages/rn-tester/js/utils/RNTesterList.android.js
@@ -31,6 +31,11 @@ const Components: Array<RNTesterModuleInfo> = [
     category: 'ListView',
     supportsTVOS: true,
   },
+  {
+    key: 'FlatList-nested',
+    module: require('../examples/FlatList/FlatList-nested'),
+    category: 'ListView',
+  },
   {
     key: 'ImageExample',
     category: 'Basic',